Progenitus stands in front of a giant Devilsaur skull on top of the tallest mountain top in Ihgaluk Crag. He can be quite tricky because you fight him hundreds of feet up above the ground level on a small platform and his abilities all involve knockback. To defeat him, you must dodge his self-centered AoE ability (denoted by a red circle around him) and his targeted missile ability (denoted by a red circle under you) or they will knock you off the cliff, resulting typically in your death or an evade. An alternative strategy is to pull Progentius from his platform and slowly work down towards the ground with him, so that even if you get knocked back, it isn't life-threatening or fight-ending.
Progentius will occasionally cast Evolution to buff himself, creating a form of soft enrage on the fight. However, the main threat comes from the ability Unleash Blood, which is a large AoE in a faint red circle around Progentius with a very low cast time. If you get hit, you will be knocked back very far and likely die or cause the fight to end. His final ability, Mutagenic Burst, is another knockback. For Mutagenic Burst, Progentius fires a missile at you (denoted by a faint red circle). Unlike Unleash Blood, I found Mutagenic Burst fairly easy to dodge even as a melee because the projectile has some travel time after the cast ends, but this ability can also ruin your day if you are not careful.
Progenitus spawns in the same place as Kroshik (egg) which is part of When in Ihgaluk, Do as the Skumblade Do - two birds with one stone.
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